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At least 253 records · Page 14

Star and Dust Formation Activities in AzTEC-3: A Starburst Galaxy at z equals 5.3

Analyses of of high-redshift ultraluminous infrared (IR) galaxies traditionally use the observed optical to submillimeter spectral energy distribution (SED) and estimates of the dynamical mass as observational constraints to derive the star formation rate (SFR), the stellar mass, and age of these objects. In this lecture we add this constraint to the analysis of AzTEC-3, a starburst galaxy at z=5.3. We construct different stellar and chemical evolutionary scenarios, constrained to produce the inferred dust mass and observed luminosity before the associated stellar mass exceeds the observational limit. A robust result of our models is that all scenarios require most of the radiating dust mass to have been accreted in molecular clouds. Our new procedure highlights the importance of a multi wavelength approach, and of the use of dust evolution models in constraining the age and the star formation activity and history in galaxies.

Dwek, Eliahu↗

Star and Dust Formation Activities in AzTEC-3: A Starburst Galaxy at z=5.3

Analyses of of high-redshift ultraluminous infrared (IR) galaxies traditionally use the observed optical to submillimeter spectral energy distribution (SED) and estimates of the dynamical mass as observational constraints to derive the star formation rate (SFR), the stellar mass, and age of these objects. In this lecture we add this constraint to the analysis of AzTEC-3, a starburst galaxy at z=5.3. We construct different stellar and chemical evolutionary scenarios, constrained to produce the inferred dust mass and observed luminosity before the associated stellar mass exceeds the observational limit. A robust result of our models is that all scenarios require most of the radiating dust mass to have been accreted in molecular clouds. Our new procedure highlights the importance of a multiwavelength approach, and of the use of dust evolution models in constraining the age and the star formation activity and history in galaxies.

Dwek, Eliahu↗

On the evolutionary status of bright, low-mass X-ray sources

A model of bright, low-mass X-ray binaries is proposed which features a lower giant-branch star losing mass on a nuclear time scale to an accreting compact companion. Simple numerical models show that mass transfer rates equal to or greater than 10 to the -9th solar masses per yr are sustained at very nearly a constant rate until the envelope of the donor star is exhausted. The model predicts orbital periods in the range 1-200 days and X-ray to optical luminosity ratios Lx/Lopt = 200-1000 for these sources. It accounts in a natural way for the large fraction of the total galactic bulge luminosity emitted by a few bright (10 to the 37th erg/s or greater) sources. It also accords very well with the observed X-ray and optical properties of the halo source Cyg X-2 and also with those of 2S 0921-63, provided this latter system contains a massive accreting white dwarf rather than a neutron star. Problems of the prior evolution of low-mass X-ray sources are also briefly delineated.

Webbink, R. F.↗

Analysis of in Vitro Evolution Reveals the Underlying Distribution of Catalytic Activity Among Random Sequences

The emergence of catalytic RNA is believed to have been a key event during the origin of life. Understanding how catalytic activity is distributed across random sequences is fundamental to estimating the probability that catalytic sequences would emerge. Here, we analyze the in vitro evolution of triphosphorylating ribozymes and translate their fitnesses into absolute estimates of catalytic activity for hundreds of ribozyme families. The analysis efficiently identified highly active ribozymes and estimated catalytic activity with good accuracy. The evolutionary dynamics follow Fisher’s Fundamental Theorem of Natural Selection and a corollary, permitting retrospective inference of the distribution of fitness and activity in the random sequence pool for the first time. The frequency distribution of rate constants appears to be log-normal, with a surprisingly steep dropoff at higher activity, consistent with a mechanism for the emergence of activity as the product of many independent contributions.

Abe Pressman↗

Unique Structural Features Relate to Evolutionary Adaptation of Cytochrome P450 in the Abyssal Zone

Cytochromes P450 (CYPs) form one of the largest enzyme superfamilies, with similar structural folds yet biological functions varying from synthesis of physiologically essential compounds to metabolism of myriad xenobiotics. Sterol 14α-demethylases (CYP51s) represent a very special P450 family, regarded as a possible evolutionary progenitor for all currently existing P450s. In metazoans CYP51 is critical for the biosynthesis of sterols including cholesterol. Here we determined the crystal structures of ligand-free CYP51s from the abyssal fish Coryphaenoides armatus and human-. Comparative sequence–structure–function analysis revealed specific structural elements that imply elevated conformational flexibility, uncovering a molecular basis for faster catalytic rates, lower substrate selectivity, and intrinsic resistance to inhibition. In addition, the C. armatus structure displayed a large-scale repositioning of structural segments that, in vivo, are immersed in the endoplasmic reticulum membrane and border the substrate entrance (the FG arm, >20 Å, and the β4 hairpin, >15 Å). The structural distinction of C. armatus CYP51, which is the first structurally characterized deep sea P450, suggests stronger involvement of the membrane environment in regulation of the enzyme function. We interpret this as a co-adaptation of the membrane protein structure with membrane lipid composition during evolutionary incursion to life in the deep sea.

Biochemistry & Molecular Biology↗

Flare loop radiative hydrodynamics. I - Basic methods

The study presented here has two goals. The first is to examine the role of energy transport mechanisms in flare dynamics, for instance, thermal cone conduction in the corona and radiative transfer in the chromosphere. The second is to provide diagnostics of flare energization processes by predicting the evolutionary behavior of concurrent emissions from all regions of the flaring atmosphere. To attain these ends, a numerical method is developed for the simultaneous solution of thy continuity, momentum, and energy equations; the time-dependent atomic rate equations describing ionization and excitation; and the radiative transfer equations. With the physical model and computational methods used here, all components of the plasma move as a single fluid and in the initial studies are assumed to have a common kinetic temperature. The plasma is constrained to move in one dimension along the axis of a loop of nonuniform cross section, the geometry of which is defined by a sufficiently strong magnetic field.

Mcclymont, A. N.↗

The frequency of T Tauri companion stars

We present the results of a magnitude-limited multiplicity survey of T Tauri stars in two nearby star-forming regions, Taurus-Auriga and Ophiuchus-Scorpius. Each of the 69 stars in the sample was observed at K with an infrared array camera on the Hale 5-m Telescope at Palomar Observatory and imaged using 2D speckle imaging techniques. Thirty-three companion stars were found of which 15 were new detections. One of the main results of this survey indicates that the binary star frequency in the projected linear separation range 14 to 225 AU for T Tauri stars is a factor of 3.5 greater than that of the solar-type main-sequence stars. Given the limited angular separation range of this survey, the rate at which binaries are detected suggests that most T Tauri stars have companions. We propose that the observed overabundance of companions to T Tauri stars relative to their older counterparts on the main sequence is an evolutionary effect; in this scheme, triple and higher order T Tauri star systems, which are observed at higher frequencies than for the solar-type main-sequence stars, are disrupted by close encounters with another star or system of stars.

Ghez, A. M.↗

A fully associative, nonisothermal, nonlinear kinematic, unified viscoplastic model for titanium alloys

Specific forms for both the Gibb's and complementary dissipation potentials are chosen such that a complete (i.e., fully associative) potential base multiaxial, nonisothermal unified viscoplastic model is obtained. This model possesses one tensorial internal state variable (that is, associated with dislocation substructure) and an evolutionary law that has nonlinear kinematic hardening and both thermal and strain induced recovery mechanisms. A unique aspect of the present model is the inclusion of nonlinear hardening through the use of a compliance operator, derived from the Gibb's potential, in the evolution law for the back stress. This nonlinear tensorial operator is significant in that it allows both the flow and evolutionary laws to be fully associative (and therefore easily integrated), greatly influences the multiaxial response under non-proportional loading paths, and in the case of nonisothermal histories, introduces an instantaneous thermal softening mechanism proportional to the rate of change in temperature. In addition to this nonlinear compliance operator, a new consistent, potential preserving, internal strain unloading criterion has been introduced to prevent abnormalities in the predicted stress-strain curves, which are present with nonlinear hardening formulations, during unloading and reversed loading of the external variables. The specific model proposed is characterized for a representative titanium alloy commonly used as the matrix material in SiC fiber reinforced composites, i.e., TIMETAL 21S. Verification of the proposed model is shown using 'specialized' non-standard isothermal and thermomechanical deformation tests.

Arnold, S. M.↗

Modeling of Thermal Decomposition of TATB-Based Explosive for Safety Analysis

We investigate and model the cook-off behavior of LX-17 to understand the response of explosive systems in abnormal thermal environments. Decomposition has been explored via conventional ODTX (One-Dimensional Time-to-eXplosion), PODTX (ODTX with pressure-measurement), TGA (Thermo-Gravimetric Analysis), and DSC (Differential Scanning Calorimetry) experiments under isothermal and ramped temperature profiles. The data were used to fit reaction rate parameters for proposed schemes in an ALE3D computational model. This model includes chemical reactions, thermo- and hydro-dynamics, and material properties, including thermal expansion, compressibility, and strength. These parameterizations were carried out utilizing a Python evolutionary optimization method on LLNL’s high-performance computing clusters. Additional experiments are being developed to further characterize and monitor decomposition intermediates to improve the model. Once experimentally validated, this model will be scalable to several applications involving LX-17. Furthermore, the optimization methodology developed herein should be applicable to other high explosive materials.

Chemistry - Chemical explosives↗

Evolutionary space station fluids management strategies

Results are summarized for an 11-month study to define fluid storage and handling strategies and requirements for various specific mission case studies and their associated design impacts on the Space Station. There are a variety of fluid users which require a variety of fluids and use rates. Also, the cryogenic propellants required for NASA's STV, Planetary, and Code Z missions are enormous. The storage methods must accommodate fluids ranging from a high pressure gas or supercritical state fluid to a sub-cooled liquid (and superfluid helium). These requirements begin in the year 1994, reach a maximum of nearly 1800 metric tons in the year 2004, and trail off to the year 2018, as currently planned. It is conceivable that the cryogenic propellant needs for the STV and/or Lunar mission models will be met by LTCSF LH2/LO2 tanksets attached to the SS truss structure. Concepts and corresponding transfer and delivery operations have been presented for STV propellant provisioning from the SS. A growth orbit maneuvering vehicle (OMV) and associated servicing capability will be required to move tanksets from delivery launch vehicles to the SS or co-orbiting platforms. Also, appropriate changes to the software used for OMV operation are necessary to allow for the combined operation of the growth OMV. To support fluid management activities at the Space Station for the experimental payloads and propellant provisioning, there must be truss structure space allocated for fluid carriers and propellant tanksets, and substantial beam strengthening may be required. The Station must have two Mobile Remote Manipulator Systems (MRMS) and the growth OMV propellant handling operations for the STV at the SS. Propellant needs for the Planetary Initiatives and Code Z mission models will most likely be provided by co-orbiting propellant platform(s). Space Station impacts for Code Z mission fluid management activities will be minimal.

Source record↗

Theoretical and experimental prerequisites of exobiology

Exobiology and the origin of life: The Inseparable Connection The world we live in is constantly changing through a process of evolutionary development that is gradual, and which leads to emergence of ever newer, more complex, more diverse, and richly differentiated forms of reality. This process may not occur simultaneously and identically everywhere, but in different ways at different rates in various places in the universe and on various objects in the celestial world. Thus, evolution must not be diagramed as a single direct line, but rather as a cluster of various paths, whose separate branches may lead to very complex and complete forms of organization and movement of matter. Nothing is known about many of these forms, and in numerous instances, their existence is not even suspected. Any one form should not be considered prematurely as a special category of life, only because of its complexity or perfection. Life is far from a result of all the numerous branches in the development of matter; its specific qualities and ways of emerging and improving are inherent. Only a single example of life, terrestrial life, is known at present, from which judgments must proceed of other possible forms of biological organization. The Earth and the events which originated on it must, to a great extent, serve as a model for broader judgments of life in the universe.

A. I. Oparin↗

Solar nebula dispersal and the stability of the planetary system. I - Scanning secular resonance theory

Secular resonances in the early solar system are investigated with the aim of establishing constraints on the time scale and method of solar nebula dispersal. Simplified nebula models and dispersal routines are used to approximate changes in an assumed axisymmetric nebula potential. These changes drive an evolutionary sequence of Laplace-Lagrange solutions for the secular variations of the solar system. These sequences are characterized by a sweep of one or more giant planet resonances through the inner solar system. Their effect is rate-dependent; characteristic dispersal times of not greater than 10,000 to 100,000 years are required to avoid the generation of terrestrial eccentricities and inclinations in excess of observed values.

Ward, W. R.↗

Genomic clocks and evolutionary timescales

For decades, molecular clocks have helped to illuminate the evolutionary timescale of life, but now genomic data pose a challenge for time estimation methods. It is unclear how to integrate data from many genes, each potentially evolving under a different model of substitution and at a different rate. Current methods can be grouped by the way the data are handled (genes considered separately or combined into a 'supergene') and the way gene-specific rate models are applied (global versus local clock). There are advantages and disadvantages to each of these approaches, and the optimal method has not yet emerged. Fortunately, time estimates inferred using many genes or proteins have greater precision and appear to be robust to different approaches.

Evolution, Molecular↗

Estimating rates and patterns of diversification with incomplete sampling: a case study in the rosids

Premise Recent advances in generating large‐scale phylogenies enable broad‐scale estimation of species diversification. These now common approaches typically are characterized by (1) incomplete species coverage without explicit sampling methodologies and/or (2) sparse backbone representation, and usually rely on presumed phylogenetic placements to account for species without molecular data. We used empirical examples to examine the effects of incomplete sampling on diversification estimation and provide constructive suggestions to ecologists and evolutionary biologists based on those results. Methods We used a supermatrix for rosids and one well‐sampled subclade (Cucurbitaceae) as empirical case studies. We compared results using these large phylogenies with those based on a previously inferred, smaller supermatrix and on a synthetic tree resource with complete taxonomic coverage. Finally, we simulated random and representative taxon sampling and explored the impact of sampling on three commonly used methods, both parametric (RPANDA and BAMM) and semiparametric (DR). Results We found that the impact of sampling on diversification estimates was idiosyncratic and often strong. Compared to full empirical sampling, representative and random sampling schemes either depressed or inflated speciation rates, depending on methods and sampling schemes. No method was entirely robust to poor sampling, but BAMM was least sensitive to moderate levels of missing taxa. Conclusions We suggest caution against uncritical modeling of missing taxa using taxonomic data for poorly sampled trees and in the use of summary backbone trees and other data sets with high representative bias, and we stress the importance of explicit sampling methodologies in macroevolutionary studies.

59 BASIC BIOLOGICAL SCIENCES↗

Stability of Cool Cores during Galaxy Cluster Growth: A Joint Chandra/SPT Analysis of 67 Galaxy Clusters along a Common Evolutionary Track Spanning 9 Gyr

We present the results of a joint analysis of Chandra X-ray and South Pole Telescope (SPT) Sunyaev–Zel’dovich (SZ) observations targeting the first sample of galaxy clusters at 0.3 < z < 1.3, selected to be the progenitors of well-studied nearby clusters based on their expected accretion rate. We develop a new procedure in order to tackle the analysis challenge that is estimating the intracluster medium (ICM) properties of low-mass and high-redshift clusters with ~150 X-ray counts. One of the dominant sources of uncertainty on the ICM density profile estimated with a standard X-ray analysis with such shallow X-ray data is due to the systematic uncertainty associated with the ICM temperature obtained through the analysis of the background-dominated X-ray spectrum. We show that we can decrease the uncertainty on the density profile by a factor varying between 2 and 8 with a joint deprojection of the X-ray surface brightness profile measured by Chandra and the SZ-integrated Compton parameter available in the SPT cluster catalog. We apply this technique to the whole sample of 67 clusters in order to track the evolution of the ICM core density during cluster growth. We confirm that the evolution of the gas density profile is well modeled by the combination of a fixed core and a self-similarly evolving non-cool-core profile. We show that the fraction of cool cores in this sample is remarkably stable with redshift although clusters have gained a factor of ~4 in total mass over the past ~9 Gyr.

79 ASTRONOMY AND ASTROPHYSICS↗

The western red cedar (Thuja plicata) 8-8' DIRIGENT family displays diverse expression patterns and conserved monolignol coupling specificity

The isolation and characterization of a multigene family of the first class of dirigent proteins (namely that mainly involved in 8-8' coupling leading to (+)-pinoresinol in this case) is reported, this comprising of nine western red cedar (Thuja plicata) DIRIGENT genes (DIR1-9) of 72-99.5% identity to each other. Their corresponding cDNA clones had coding regions for 180-183 amino acids with each having a predicted molecular mass of ca. 20 kDa including the signal peptide. Real time-PCR established that the DIRIGENT isovariants were differentially expressed during growth and development of T. plicata (P < 0.05). The phylogenetic relationships and the rates and patterns of nucleotide substitution suggest that the DIRIGENT gene may have evolved via paralogous expansion at an early stage of vascular plant diversification. Thereafter, western red cedar paralogues have maintained an high homogeneity presumably via a concerted evolutionary mode. This, in turn, is assumed to be the driving force for the differential formation of 8-8'-linked pinoresinol derived (poly)lignans in the needles, stems, bark and branches, as well as for massive accumulation of 8-8'-linked plicatic acid-derived (poly)lignans in heartwood.

Non-NASA Center↗

The evolutionary thermal response of a white dwarf to compressional heating by periodic dwarf nova accretion events

The thermal response of the underlying white dwarf in a cataclysmic variable to the periodic deposition of mass by a dwarf nova accretion event is simulated with a quasi-static stellar evolution code incorporating time variable accretion. After accretion at rates typical of dwarf nova outbursts (approx. 10(exp -7) to 10(exp -8)/yr) for outburst durations of days to 2 weeks, the radial infall is shut off and the evolution of the white dwarf is followed during dwarf nova quiescence. The matter is assumed to accrete softly with the same entropy as the white dwarf outer layers. In some sequences accretion is resumed and shut off repeatedly at intervals of months to simulate the thermal evolution of the white dwarf in typical dwarf novae such as SS Cygni and U Geminorum, between successive dwarf nova outbursts. Thermal timescales for white dwarf cooling following a given dwarf nova outburst depend upon the accretion rate, outburst duration, and white dwarf mass; they are nominally in the range 0.2-0.8 years for parameters typical of dwarf novae (white dwarf masses in the range 1.2-0.6 solar mass, outburst accretion rates in the range 1 x 10(exp -7)-10(exp -8) solar mass/yr, outburst durations in the range 7-14, days and quiescent intervals of 30-300 days). If the e-folding timescale of the white dwarf cooling is shorter than the quiescent interval bewteen outbursts, then the effect of compressional heating is too small to be observationally detectable.

Sion, Edward M.↗

Spectral evolution of a subclass of gamma-ray bursts observed by batse

Among the gamma-ray bursts (GRBs) observed by the Burst and Transient Source Experiment (BATSE) on board the Compton Gamma Ray Observatory we define a subclass of bursts based on similar morphology: a sharp rise followed by a longer decay time. About 7% of all the gamma-ray bursts observed by BATSE fall into this subclass. We study the spectral evolution of these bursts by fitting models to time-segmented burst spectra and find no clear distinction between the spectral evolutionary properties of this subclass and those of other bursts. Further, we study the high time resolution spectral evolution of this subclass of GRBs using their spectral hardness ratios. A majority of the bursts show hardness ratio leading the counting rate and also display a continuous hard to soft evolution. The time lag between the counting rate and the hardness ratio is found to be directly correlated with the rise time of the counting rate profile. We also find, for the first time, evidence for spectral variation in a timescale of 64 ms.

Bhat, P. N.↗