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Search indexed NASA NTRS and DOE OSTI research on propulsion, heat transfer, battery materials and energy systems. Follow report and document links to the original sources.

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All sourcesNASA NTRSDOE OSTI
NASA NTRS1993

The analysis of global cloud and radiation data for the study of cloud-climate interactions

During the past year, ISCCP C1 data has been analyzed for two regions, the stratocumulus area off California and the tropical Atlantic. The first part of the study dealing with stratocumulus clouds is complete and the results form the technical portion of this report. A manuscript that has recently been prepared covering this subject is attached as an Appendix. The abstract describes the key finding that cloudy mean liquid water path (LWP) is invariant with cloud fraction for cloud cover between 20% - 80%. Similar conclusions are reached with ISCCP and LANDSAT data. This has important implications for General Circulation Models. A possible application is described in the Discussion section of the Appendix. An idea that has been proposed for dealing with GCM hydrology can be used.

HARSHVARDHAN↗
NASA NTRS1993

Cloud properties from the analysis of AVHRR observations for FIRE 2

Preliminary results are presented for cloud properties from the analysis of AVHRR observations for FIRE 2. The properties were obtained from a combination of the spatial coherence method and a multispectral retrieval scheme. Geographically gritted fields for the number of cloud layers were produced. For single layered cloud systems, fractional cloud cover, cloud emission temperature, cloud emissivity, and particle size were retrieved. Statistics on the properties of upper-level clouds and the Coffeeville cloud conditions are presented.

Lin, Xijian↗
NASA NTRS2007

An Improved Algorithm for Retrieving Surface Downwelling Longwave Radiation from Satellite Measurements

Zhou and Cess [2001] developed an algorithm for retrieving surface downwelling longwave radiation (SDLW) based upon detailed studies using radiative transfer model calculations and surface radiometric measurements. Their algorithm linked clear sky SDLW with surface upwelling longwave flux and column precipitable water vapor. For cloudy sky cases, they used cloud liquid water path as an additional parameter to account for the effects of clouds. Despite the simplicity of their algorithm, it performed very well for most geographical regions except for those regions where the atmospheric conditions near the surface tend to be extremely cold and dry. Systematic errors were also found for scenes that were covered with ice clouds. An improved version of the algorithm prevents the large errors in the SDLW at low water vapor amounts by taking into account that under such conditions the SDLW and water vapor amount are nearly linear in their relationship. The new algorithm also utilizes cloud fraction and cloud liquid and ice water paths available from the Cloud and the Earth's Radiant Energy System (CERES) single scanner footprint (SSF) product to separately compute the clear and cloudy portions of the fluxes. The new algorithm has been validated against surface measurements at 29 stations around the globe for Terra and Aqua satellites. The results show significant improvement over the original version. The revised Zhou-Cess algorithm is also slightly better or comparable to more sophisticated algorithms currently implemented in the CERES processing and will be incorporated as one of the CERES empirical surface radiation algorithms.

Zhou, Yaping↗
NASA NTRS2007

What You Need to Know About the OMI NO2 Data Product for Air Quality Studies

The standard nitrogen dioxide (NO2) data product, produced from measurements by the Ozone Monitoring Instrument (OMI), are publicly available online from the NASA GESDISC facility. Important data fields include total and tropospheric column densities, as well as collocated data for cloud fraction and cloud top height, surface albedo and snow/ice coverage, at the resolution of the OMI instrument (12 km x 26 km, at nadir). The retrieved NO2 data have been validated, principally under clear-sky conditions. The first public-release version has been available since September 2006. An improved version of the data product, which includes a number of new data fields, and improved estimates of the retrieval uncertainties will be released by the end of 2007. This talk will describe the standard NO2 data product, including details that are essential for the use of the data for air quality studies. We will also describe the principal improvements with the new version of the data product.

Celarier, E. A.↗
NASA NTRS2006

An Improved Algorithm for Retrieving Surface Downwelling Longwave Radiation from Satellite Measurements

Retrieving surface longwave radiation from space has been a difficult task since the surface downwelling longwave radiation (SDLW) are integrations from radiation emitted by the entire atmosphere, while those emitted from the upper atmosphere are absorbed before reaching the surface. It is particularly problematic when thick clouds are present since thick clouds will virtually block all the longwave radiation from above, while satellites observe atmosphere emissions mostly from above the clouds. Zhou and Cess developed an algorithm for retrieving SDLW based upon detailed studies using radiative transfer model calculations and surface radiometric measurements. Their algorithm linked clear sky SDLW with surface upwelling longwave flux and column precipitable water vapor. For cloudy sky cases, they used cloud liquid water path as an additional parameter to account for the effects of clouds. Despite the simplicity of their algorithm, it performed very well for most geographical regions except for those regions where the atmospheric conditions near the surface tend to be extremely cold and dry. Systematic errors were also found for areas that were covered with ice clouds. An improved version of the algorithm was developed that prevents the large errors in the SDLW at low water vapor amounts. The new algorithm also utilizes cloud fraction and cloud liquid and ice water paths measured from the Cloud and the Earth's Radiant Energy System (CERES) satellites to separately compute the clear and cloudy portions of the fluxes. The new algorithm has been validated against surface measurements at 29 stations around the globe for the Terra and Aqua satellites. The results show significant improvement over the original version. The revised Zhou-Cess algorithm is also slightly better or comparable to more sophisticated algorithms currently implemented in the CERES processing. It will be incorporated in the CERES project as one of the empirical surface radiation algorithms.

Zhou, Yaping↗
NASA NTRS2014

Merging of OMI and AIRS Ozone Data

The OMI Instrument measures ozone using the backscattered light in the UV part of the spectrum. In polar night there are no OMI measurements so we hope to incorporate the AIRS ozone data to fill in these missing regions. AIRS is on the Aqua platform and has been operating since May 2002. AIRS is a multi-detector array grating spectrometer containing 2378 IR channels between 650 per centimeter and 2760 per centimeter which measures atmospheric temperature, precipitable water, water vapor, CO, CH4, CO2 and ozone profiles and column amount. It can also measure effective cloud fraction and cloud top pressure for up to two cloud layers and sea-land skin temperature. Since 2008, OMI has had part of its aperture occulted with a piece of the thermal blanket resulting in several scan positions being unusable. We hope to use the AIRS data to fill in the missing ozone values for those missing scan positions.

OMI↗
NASA NTRS2014

Ticosonde CFH at Costa Rica: A Seasonal Climatology of Tropical UT-LS Water Vapor and Inter-Comparisons with MLS and CALIPSO

Balloon sonde measurements of tropical water vapor using the Cryogenic Frostpoint Hygrometer were initiated in Costa Rica in July 2005 and have continued to the present day. Over the nine years through July 2014, the Ticosonde program has launched 174 CFH payloads, representing the longest-running and most extensive single-site balloon dataset for tropical water vapor. In this presentation we present a seasonal climatology for water vapor and ozone at Costa Rica and examine the frequency of upper tropospheric supersaturation with comparisons to cloud fraction and cloud ice water content observations from the Cloud Aerosol Lidar with Orthogonal Polarization (CALIOP) on the CALIPSO mission. We then make a critical comparison of these data to water vapor measurements from the MLS instrument on board Aura in light of recently published work for other sites. Finally, we examine time series of 2-km altitude averages in the upper troposphere-lower stratosphere at Costa Rica in light of anomalies and trends seen in various large-scale indices of tropical water vapor.

water vapor↗
NASA NTRS2016

Impact of MODIS SWIR Band Calibration Improvements on Level-3 Atmospheric Products

The spectral reflectance measured by the MODIS reflective solar bands (RSB) is used for retrieving many atmospheric science products. The accuracy of these products depends on the accuracy of the calibration of the RSB. To this end, the RSB of the MODIS instruments are primarily calibrated on-orbit using regular solar diffuser (SD) observations. For lambda < 0.94 microns the SDs on-orbit bi-directional reflectance factor (BRF) change is tracked using solar diffuser stability monitor (SDSM) observations. For lambda > 0.94 microns, the MODIS Characterization Support Team (MCST) developed, in MODIS Collection 6 (C6), a time-dependent correction using observations from pseudo-invariant earth-scene targets. This correction has been implemented in C6 for the Terra MODIS 1.24 micron band over the entire mission, and for the 1.375 micron band in the forward processing. As the instruments continue to operate beyond their design lifetime of six years, a similar correction is planned for other short-wave infrared (SWIR) bands as well. MODIS SWIR bands are used in deriving atmosphere products, including aerosol optical thickness, atmospheric total column water vapor, cloud fraction and cloud optical depth. The SD degradation correction in Terra bands 5 and 26 impact the spectral radiance and therefore the retrieval of these atmosphere products. Here, we describe the corrections to Bands 5 (1.24 microns) and 26 (1.375 microns), and produce three sets (B5, B26 correction on/on, on/off, and off/off) of Terra-MODIS Level 1B (calibrated radiance product) data. By comparing products derived from these corrected and uncorrected Terra MODIS Level 1B (L1B) calibrations, dozens of L3 atmosphere products are surveyed for changes caused by the corrections, and representative results are presented. Aerosol and water vapor products show only small local changes, while some cloud products can change locally by > 10%, which is a large change.

Calibration↗
NASA NTRS2020

Differences in Tropical High Clouds Among Reanalyses: Origins and Radiative Impacts

We examine differences among reanalysis highcloud products in the tropics, assess the impacts of these differences on radiation budgets at the top of the atmosphere and within the tropical upper troposphere and lower stratosphere (UTLS), and discuss their possible origins in the context of the reanalysis models. We focus on the ERA5 (fifthgeneration European Centre for Medium-range Weather Forecasts – ECMWF – reanalysis), ERA-Interim (ECMWF Interim Reanalysis), JRA-55 (Japanese 55-year Reanalysis), MERRA-2 (Modern-Era Retrospective Analysis for Research and Applications, Version 2), and CFSR/CFSv2 (Climate Forecast System Reanalysis/Climate Forecast System Version 2) reanalyses. As a general rule, JRA-55 produces the smallest tropical high-cloud fractions and cloud water contents among the reanalyses, while MERRA-2 produces the largest. Accordingly, long-wave cloud radiative effects are relatively weak in JRA-55 and relatively strong in MERRA-2. Only MERRA-2 and ERA5 among the reanalyses produce tropical-mean values of outgoing long-wave radiation (OLR) close to those observed, but ERA5 tends to underestimate cloud effects, while MERRA-2 tends to overestimate variability. ERA5 also produces distributions of longwave, short-wave, and total cloud radiative effects at the top of the atmosphere that are very consistent with those observed. The other reanalyses all exhibit substantial biases in at least one of these metrics, although compensation between the long-wave and short-wave effects helps to constrain biases in the total cloud radiative effect for most reanalyses. The vertical distribution of cloud water content emerges as a key difference between ERA-Interim and other reanalyses. Whereas ERA-Interim shows a monotonic decrease of cloud water content with increasing height, the other reanalyses all produce distinct anvil layers. The latter is in better agreement with observations and yields very different profiles of radiative heating in the UTLS. For example, whereas the altitude of the level of zero net radiative heating tends to be lower in convective regions than in the rest of the tropics in ERAInterim, the opposite is true for the other four reanalyses. Differences in cloud water content also help to explain systematic differences in radiative heating in the tropical lower stratosphere among the reanalyses. We discuss several ways in which aspects of the cloud and convection schemes impact the tropical environment. Discrepancies in the vertical profiles of temperature and specific humidity in convective regions are particularly noteworthy, as these variables are directly constrained by data assimilation, are widely used, and feed back to convective behaviour through their relationships with thermodynamic stability.

High Clouds↗
NASA NTRSDate not supplied

Examining the Feasibility of Detailed Decomposition of Radiative Flux Anomalies to Cloud Changes

This presentation will explore the degree to which radiative flux anomalies in the last two decades seen by CERES can be interpreted as the result of changes in particular cloud types. A study of this kind is potentially enabled by the new CERES FlxByCldTyp (FBCT) product which provides combined Terra and Aqua daytime 1°-regional gridded daily and monthly top-of-the-atmosphere radiative fluxes and associated MODIS-derived cloud properties stratified by Cloud Top Pressure (CTP) and Cloud Optical Thickness (TAU), i.e., arrays of cloud properties and radiative fluxes resolved in CTP-TAU bins. It has been shown (Sun et al. 2022) that the FBCT time series of global flux anomalies tracks closely its SSF1deg and EBAF counterparts, indicating that these three products provide fluxes of consistent stability for interannual variability studies. We are thus in position to examine whether CERES regional or global radiative fluxes anomalies and trends are driven by changes in certain elements of FBCT flux arrays corresponding to particular cloud types. Anomalies in flux array elements will in turn be investigated in terms of cloud property variations within the CTP-TAU bins where the biggest flux anomalies are encountered. We will essentially seek to obtain the relative contributions to the anomalies of binned all-sky fluxes of changes in cloud fraction within cloud types and of changes in their overcast fluxes modulated by other cloud properties. Such an analysis holds great promise of unveiling the root causes of cloud-driven changes in the planetary radiation budget of the last two decades.

cloud↗
NASA NTRS2023

A Cloud Detection Neural Network for Above-Aircraft Clouds Using Airborne Cameras

We introduce a method convolutional neural networks to detect the presence of clouds in airborne camera images. We quantify the performance of this Cloud Detection Neural Network (CDNN) using human-labeled validation data where we report a 96% accuracy in detecting clouds in testing datasets for both Zenith- viewing and Forward-viewing models. We assess our performance by comparing the flight-averaged cloud fraction of zenith and forward CDNN retrievals, with that of the prototype hyperspectral total-diffuse Sunshine Pyranometer (SPN-S) instrument’s cloud optical depth data. Comparison of the CDNN with the SPN-S on time specific intervals resulted in 93% accuracy for the zenith- viewing CDNN and 84% for the forward- viewing CDNN. The comparison of the CDNNs with the SPN-S on flight-averaged cloud fraction resulted in an agreement of 0.15 for the Forward CDNN and 0.07 for the Zenith CDNN. We then quantify the ability of the CDNN to identify the presence of clouds above the aircraft using a forward- looking camera mounted inside the aircraft cockpit compared to the use of an All- Sky upward-looking camera that is mounted outside the fuselage on top of the aircraft. We present results from the CDNN based on airborne imagery from the NASA Aerosol Cloud Meteorology Interactions Over the Western Atlantic Experiment (ACTIVATE) and the Clouds, Aerosol and Monsoon Processes- Philippines Experiment (CAMP2Ex). For CAMP2Ex 53% of flight dates had above- aircraft cloud fraction above 50%, while for ACTIVATE 52% and 54% of flight dates observed above-aircraft cloud fraction above 50% for 2020 and 2021, respectively.

Joseph D Nied↗
NASA NTRSDate not supplied

Role of Arctic Low Clouds in the Arctic Climate System: Isolating the Surface Type Influence on Arctic Low-Clouds

Interactions between sea ice and clouds represent a mechanism through which sea ice influences climate. Understanding the cloud response to the rapidly changing Arctic surface properties is necessary for improving climate simulations and projection of future change inside and outside the Arctic. Towards the goal of understanding the cloud response to sea ice, we composite cloud properties for ice-free, marginal ice zone (MIZ), and ice-covered surfaces during MIZ crossing events to analyze the cloud property differences between surface types. Restricting the analysis to MIZ crossing events enables the isolation of the sea ice effect on clouds from meteorological factors. We find larger cloud fraction and total water concentration below ~1.5 km over ice-free relative to ice-covered surfaces during non-summer months. During summer, the results suggest larger cloud fraction and total water concentration over ice-free surfaces, however differences do not exceed observational uncertainty. Cloud property differences are linked to atmospheric thermodynamic profile differences, namely ice-free surfaces are warmer, moister, less stable, and have more positive surface turbulent fluxes than ice-covered surfaces. Ice-free minus ice-covered cloud property differences scale with surface temperature differences and are only found in the presence of a surface temperature difference. Our results suggest a 0.02 cloud fraction and 0.005 g m 3 total water concentration increase (~5%) at the level of maximum cloud fraction between 2000-2021 due to the observed Arctic sea ice decline in fall, corresponding to ~2 Wm 2 increase in the net surface radiative flux supporting a positive sea ice-cloud radiative feedback in fall and winter and a negative sea ice-cloud radiative feedback in spring. We propose an updated conceptual model where the average surface type influence on cloud properties is mediated by surface temperature differences between ice-free and ice-covered surfaces.

Arctic climate↗
DOE OSTI2021

Quantifying and attributing time step sensitivities in present-day climate simulations conducted with EAMv1

Abstract. This study assesses the relative importance of time integration error in present-day climate simulations conducted with the atmosphere component of the Energy Exascale Earth System Model version 1 (EAMv1) at 1∘ horizontal resolution. We show that a factor-of-6 reduction of time step size in all major parts of the model leads to significant changes in the long-term mean climate. Examples of changes in 10-year mean zonal averages include the following: up to 0.5 K of warming in the lower troposphere and cooling in the tropical and subtropical upper troposphere, 1 %–10 % decreases in relative humidity throughout the troposphere, and 10 %–20 % decreases in cloud fraction in the upper troposphere and decreases exceeding 20 % in the subtropical lower troposphere. In terms of the 10-year mean geographical distribution, systematic decreases of 20 %–50 % are seen in total cloud cover and cloud radiative effects in the subtropics. These changes imply that the reduction of temporal truncation errors leads to a notable although unsurprising degradation of agreement between the simulated and observed present-day climate; to regain optimal climate fidelity in the absence of those truncation errors, the model would require retuning. A coarse-grained attribution of the time step sensitivities is carried out by shortening time steps used in various components of EAM or by revising the numerical coupling between some processes. Our analysis leads to the finding that the marked decreases in the subtropical low-cloud fraction and total cloud radiative effect are caused not by the step size used for the collectively subcycled turbulence, shallow convection, and stratiform cloud macrophysics and microphysics parameterizations but rather by the step sizes used outside those subcycles. Further analysis suggests that the coupling frequency between the subcycles and the rest of EAM significantly affects the subtropical marine stratocumulus decks, while deep convection has significant impacts on trade cumulus. The step size of the cloud macrophysics and microphysics subcycle itself appears to have a primary impact on cloud fraction in the upper troposphere and also in the midlatitude near-surface layers. Impacts of step sizes used by the dynamical core and the radiation parameterization appear to be relatively small. These results provide useful clues for future studies aiming at understanding and addressing the root causes of sensitivities to time step sizes and process coupling frequencies in EAM. While this study focuses on EAMv1 and the conclusions are likely model-specific, the presented experimentation strategy has general value for weather and climate model development, as the methodology can help researchers identify and understand sources of time integration error in sophisticated multi-component models.

54 ENVIRONMENTAL SCIENCES↗
NASA NTRS2023

Comparison of Clouds and Cloud Feedback between AMIP5 and AMIP6

We examine the changes in clouds and cloud feedback between Phase 5 (AMIP5) and Phase 6 (AMIP6) of the Atmospheric Model Intercomparison Project. Each model is perturbed by uniformly increasing the sea surface temperature by 4 K. The simulated cloud fraction, the perturbed states and cloud radiative kernels are used to derive cloud feedback in the shortwave (SW), longwave (LW) and their sum (Net). Compared to AMIP5, the cloud fraction in AMIP6 increases by 9.1%, while the perturbation leads to a 0.25% decrease. The Net cloud feedback at the top of the atmosphere (TOA) is almost double (174%). Statistical tests support that this change is mainly due to an increase in the surface SW cloud feedback caused by optically thick, middle and low clouds. The contribution of the atmospheric Net component (12%) stems from the increase in the atmospheric LW cloud feedback, likely to play a role in weakening (strengthening) the northward (southward) meridional atmospheric energy transport, while the opposite is true for the surface LW and Net cloud feedback in the meridional oceanic energy transport. The substantial increase in cloud feedback at the TOA primarily contributes to the higher climate sensitivity. The cloud feedback spread in AMIP6 is comparable to that in AMIP5.

Clouds↗
NASA NTRS2022

Assessment of Two Stochastic Cloud Subcolumn Generators Using Observed Fields of Vertically Resolved Cloud Extinction

We evaluate two stochastic subcolumn generators used in GCMs to emulate subgrid cloud variability enabling comparisons with satellite observations and simulations of certain physical processes. Our evaluation necessitated the creation of a reference observational dataset that resolves horizontal and vertical cloud variability. The dataset combines two CloudSat cloud products that resolve two-dimensional cloud optical depth variability of liquid, ice, and mixed phase clouds when blended at ~200 m vertical and ~ 2 km horizontal scales. Upon segmenting the dataset to individual “scenes”, mean profiles of the cloud fields are passed as input to generators that produce scene-level cloud subgrid variability. The assessment of generator performance at the scale of individual scenes and in a mean sense is largely based on inferred joint histograms that partition cloud fraction within predetermined combinations of cloud top pressure –cloud optical thickness ranges. Our main finding is that both generators tend to underestimate optically thin clouds, while one of them also tends to overestimate some cloud types of moderate and high optical thickness. Associated radiative flux errors are also calculated by applying a simple transformation to the cloud fraction histogram errors, and are found to approach values almost as high as 3 Wm-2 for the cloud radiative effect in the shortwave part of the spectrum.

cloud variability↗
DOE OSTI2020

The Contributions of Shear and Turbulence to Cloud Overlap for Cumulus Clouds

The causes of vertical cloud overlap, the ratio between the cloud fraction by area and the cloud fraction by volume, are studied for fields of cumulus convection. Using large-eddy simulations, the structure of individual clouds within the cloud field is inspected and connected to the overall cloud cover of the entire cloud field. First, it is shown that the cloud overlap of the entire field can be well approximated by the total cloud overlap of the individual clouds. It is then shown that a simple maximum overlap assumption, which is commonly used for cloud fields with small layer depth, can only account for roughly half of the overlap inefficiency. This is due to the fact that maximum overlap only represents variations in cloud width as a function of depth but does not take other factors into account, most notably shear or turbulence. The effects of turbulence and wind shear on the overlap of individual clouds are first estimated as a function of cloud depth, which closes the discrepancy between observed cloud overlap and the maximum overlap assumption. Then, an empirical model is developed for the shear and turbulence contributions on overlap as a function of cloud depth and of synoptic conditions. Finally, it is shown that our model is capable of accurately predicting overlap for a large number of cloud fields and under a variety of meteorological conditions.

54 ENVIRONMENTAL SCIENCES↗
NASA NTRS1976

A single field of view method for retrieving tropospheric temperature profiles from cloud-contaminated radiance data

An iterative method is presented to retrieve single field of view (FOV) tropospheric temperature profiles directly from cloud-contaminated radiance data. A well-defined temperature profile may be calculated from the radiative transfer equation (RTE) for a partly cloudy atmosphere when the average fractional cloud amount and cloud-top height for the FOV are known. A cloud model is formulated to calculate the fractional cloud amount from an estimated cloud-top height. The method is then examined through use of simulated radiance data calculated through vertical integration of the RTE for a partly cloudy atmosphere using known values of cloud-top height(s) and fractional cloud amount(s). Temperature profiles are retrieved from the simulated data assuming various errors in the cloud parameters. Temperature profiles are retrieved from NOAA-4 satellite-measured radiance data obtained over an area dominated by an active cold front and with considerable cloud cover and compared with radiosonde data. The effects of using various guessed profiles and the number of iterations are considered.

Hodges, D. B.↗
NASA NTRS2016

Saharan Dust as a Causal Factor of Significant Cloud Cover Along the Saharan Air Layer in the Atlantic Ocean

The tropical Atlantic is frequently affected by Saharan dust intrusions. Based on MODIS cloud fraction (CF) data during the ten-year study period, we found that these dust intrusions contribute to significant cloud cover along the Saharan Air Layer (SAL). Below the temperature inversion at the SAL's base, the presence of large amounts of settling dust particles, together with marine aerosols, produces meteorological conditions suitable for the formation of shallow stratocumulus clouds. The significant cloud fraction along the SAL together with clouds over the Atlantic Inter-tropical Convergence Zone contributes to the 20% hemispheric CF asymmetry between the tropical North and South Atlantic. This leads to the imbalance in strong solar radiation, which reaches the sea surface between the tropical North and South Atlantic, and, consequently, affects climate formation in the tropical Atlantic. Therefore, despite the fact that, over the global ocean, there is no noticeable hemispheric asymmetry in cloud fraction, over the significant area such as the tropical Atlantic the hemispheric asymmetry in CF takes place. Saharan dust is also the major contributor to hemispheric aerosol asymmetry over the tropical Atlantic. The NASA GEOS-5 model with aerosol data assimilation was used to extend the MERRA reanalysis with five atmospheric aerosol species (desert dust, sulfates, organic carbon, black carbon, and sea-salt). The obtained ten-year (2002 - 2012) MERRA-driven aerosol reanalysis dataset (aka MERRAero) showed that, over the tropical Atlantic, dust and carbonaceous aerosols were distributed asymmetrically relative to the equator, while other aerosol species were distributed more symmetrically.

CF↗