Incubation temperature as a constraint on clutch size evolution
Not provided.
Environmental Sciences & Ecology↗
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Method and apparatus for executing an asynchronous clutch-to-clutch shift in a hybrid transmission
A hybrid transmission includes first and second electric machines. A method for operating the hybrid transmission in response to a command to execute a shift from an initial continuously variable mode to a target continuously variable mode includes increasing torque of an oncoming clutch associated with operating in the target continuously variable mode and correspondingly decreasing a torque of an off-going clutch associated with operating in the initial continuously variable mode. Upon deactivation of the off-going clutch, torque outputs of the first and second electric machines and the torque of the oncoming clutch are controlled to synchronize the oncoming clutch. Upon synchronization of the oncoming clutch, the torque for the oncoming clutch is increased and the transmission is operated in the target continuously variable mode.
42 ENGINEERING↗
Free-moving artificial eggs containing temperature loggers reveal remarkable within-clutch variance in incubation temperature
Incubation is a crucial aspect of avian parental care and measuring incubation temperature in the wild can improve our understanding of life history tradeoffs and inform conservation efforts. However, there are challenges associated with measuring the temperature of eggs in natural nests. Most studies to date have measured incubation temperature by using a single, stationary logger in each nest. However, real eggs are rotated and moved throughout the nest by the parent during the incubation period, and thus, a stationary logger may not accurately represent the temperature experienced by individual eggs within the entire clutch. We recorded incubation temperature in nests by using multiple, mobile artificial egg temperature loggers. We installed six mobile loggers and one stationary logger in wood duck Aix sponsa nests to compare the two logger types in the field. We found that at a given ambient temperature, mobile loggers recorded lower average and more variable temperatures than stationary loggers. Further, temperatures recorded by stationary loggers showed no relationship with clutch size, while mobile loggers captured temperatures that were lower and more variable as clutch size increased. Also, the multiple mobile loggers revealed that eggs within a nest experienced a substantial range of temperatures throughout the incubation period. Here, we discuss potential limitations of this method, but believe that it is a promising way to collect biologically-relevant incubation temperature data and provides an opportunity to advance our understanding of incubation temperature as a parental effect.
59 BASIC BIOLOGICAL SCIENCES↗
Male body size predicts reproductive success but not within-clutch paternity patterns in gopher tortoises ( Gopherus polyphemus )
In many vertebrates, body size is an important driver of variation in male reproductive success. Larger, more fit individuals are more likely to dominate mating opportunities, skewing siring success and resulting in lower effective population sizes and genetic diversity. The mating system of the gopher tortoise (Gopherus polyphemus) has been characterized as both female defense and scramble-competition polygyny. Mating systems are typically not fixed and can be influenced by factors such as population density, demographic structure, and environmental conditions;, however, most populations will have a predominant strategy that results from local conditions. We assessed how male body size influences patterns of paternity and reproductive success in a natural population of gopher tortoises in Florida, USA. Using microsatellites, we assigned parentage of 220 hatchlings from 31 nests collected during two reproductive seasons. Larger males were significantly more likely to sire offspring and sired more offspring than smaller males; however, the likelihood of a clutch being multiply-sired was unrelated to male body size. We also found evidence of mate fidelity across years. Although paternity patterns in this high-density population are more consistent with defense polygyny, female monopoly by males was incomplete, with both large and small males contributing to multiply-sired clutches. Additional behavioral data are needed to clarify the role of female mate selection in paternity outcomes. The context-dependence of mating systems underscores the need to compare parentage patterns across populations and to recognize the potential for more than one strategy to be employed within a single population.
59 BASIC BIOLOGICAL SCIENCES↗
Sensitivity Calculations for Systems with Polyethylene Reflector Materials Using CLUTCH
The SCALE 6.2.4 code package contains four sequences for calculating $k_{eff}$ sensitivity coefficients. Two of these sequences use deterministic transport solvers: a one-dimensional (1D) capability based on XSDRN, and a two-dimensional (2D) capability based on NEWT. These sequences are restricted to the multigroup (MG) treatment of neutron energy. The three-dimensional (3D) sequences use the KENO V.a or KENO-VI Monte Carlo transport codes and can be used to calculate sensitivity coefficients with either MG or continuous-energy (CE) transport. The 3D sensitivities are ultimately reported in an MG structure, regardless of the method used in the transport calculations. If desired, the sensitivity coefficients can be reported with very fine energy resolution from a CE calculation, but they are calculated only in the MG library structure in the MG mode. CE TSUNAMI methods are available in SCALE starting in SCALE version 6.2. Sensitivity coefficients were generated using the 3D sequences as part of the generation of the SCALE 6.2.2 Validation Report; difficulties encountered when using the CLUTCH method for thick, fissionable-material reflectors were discussed and investigated as documented in a previous paper. This paper discusses the difficulties encountered in generating accurate sensitivity coefficients using the CLUTCH technique for polyethylene reflectors for two fast spectrum benchmarks. Direct perturbation (DP) calculations were performed to confirm the accuracy of the total sensitivity coefficient for important isotopes with large sensitivities in the system. Discrepancies were detected for CLUTCH-calculated sensitivity coefficients in the reflector of a critical experiment with a radial polyethylene reflector. A simple polyethylene-reflected plutonium sphere was then used to further investigate the discrepancy. Calculations performed using the iterated fission probability (IFP) method generated accurate sensitivity coefficients in both cases. The results of this study emphasize the need to confirm CLUTCH sensitivity results with DP calculations. IFP calculations are generally less efficient but more reliable than CLUTCH calculations. Improvements to the CLUTCH methodology that retain the greater efficiency but address identified difficulties are therefore potentially useful to analysts.
73 NUCLEAR PHYSICS AND RADIATION PHYSICS↗
Sex ratio of Western Bluebirds ( Sialia mexicana ) is mediated by phenology and clutch size
Mothers may produce more of one sex to maximize their fitness if there are differences in the cost of producing each sex or there are differences in their relative reproductive value. Breeding date and clutch size are known to influence offspring sex ratios in birds through sex differences in dispersal, social behaviours, differential mortality and available food resources. We tested whether breeding date, clutch size and drought conditions influenced offspring sex ratios in a sexually size-monomorphic species, the Western Bluebird Sialia mexicana, by interrogating a 21-year dataset. After controlling for differential mortality, we found that hatch dates late in the breeding season were associated with the production of more females, suggesting that the value of producing males declines as the breeding season progresses. When clutch size was taken into account, small clutches yielded significantly more females late in the breeding season than in the early and middle parts of the breeding season, which produced significantly more males. Here, large clutches early in the season tended to produce more females, although this was not significant. Drought severity was not correlated with sex ratio adjustment. We propose and discuss several explanations for these patterns, including male offspring, but not female offspring, acting as helpers, increased female nestling provisioning late in the breeding season, differences in food abundance and egg-laying order. Future work will help to uncover the mechanisms leading to these patterns. Identifying patterns and mechanisms of sex ratio skew from long-term datasets is important for informing predictions regarding life-history trade-offs in wildlife populations.
59 BASIC BIOLOGICAL SCIENCES↗
Method and apparatus for operating a powertrain system upon detecting a stuck-closed clutch
A powertrain system includes a multi-mode transmission having a plurality of torque machines. A method for controlling the powertrain system includes identifying all presently applied clutches including commanded applied clutches and the stuck-closed clutch upon detecting one of the torque-transfer clutches is in a stuck-closed condition. A closed-loop control system is employed to control operation of the multi-mode transmission accounting for all the presently applied clutches.
42 ENGINEERING↗
Egg and clutch sizes of western chicken turtles (Deirochelys reticularia miaria)
Chicken turtles (Deirochelys reticularia) are generally characterized as having atypical reproductive characteristics relative to other sympatric emydids. However, the comparatively understudied western chicken turtle (D. r. miaria) has been shown to exhibit some reproductive characteristics that differ from the other subspecies. Therefore, we examined clutch and egg sizes from six D. r. miaria (13 clutches) in Oklahoma and compared the results to values that have been reported for the other two subspecies. Females nested up to three times per year, with clutches ranging from 8-13 eggs per clutch (mean = 10.9). The eggs were 19.4-25.3 mm wide (mean = 22.2 mm). These values are greater than the means reported for the other subspecies, but the differences were not statistically significant.
59 BASIC BIOLOGICAL SCIENCES↗
Adapting CLUTCH methodology to multigroup TSUNAMI-3D for eigenvalue sensitivity calculations
The sensitivity of the eigenvalue to uncertainties in nuclear data and its evaluation are important for nuclear criticality safety. TSUNAMI-3D sequences within the SCALE code system offer several options to the user community for calculating eigenvalue sensitivity coefficients with multigroup (MG) and continuous energy (CE) 3D transport capabilities. TSUNAMI-3D sequences implement the adjoint-based perturbation theory with MG KENO code, the Contributon Linked eigenvalue sensitivity/Uncertainty estimation via Track length importance CHaracterization (CLUTCH) method with CE KENO code, and the Iterated Fission Probability (IFP) method with CE KENO and Shift codes. Each method has benefits and limitations depending on the problem that is run. The work presented here aims to adapt the CLUTCH method, which enables the Contributon method's mesh-free, memory-efficient approach for calculating adjoint-weighted tallies for sensitivity calculations, to the MG TSUNAMI-3D sequence. This application would eliminate the explicit adjoint KENO calculation, as well as the memory-consuming mesh flux moment tallies required by the conventional MG TSUNAMI-3D. Smaller memory footprints in the CLUTCH methodology and relatively shorter runtimes in MG KENO transport can make MG TSUNAMI-3D a viable method for some complex problems. Moreover, this adaptation allows MG sensitivity calculations with Shift, ORNL's next-generation high-performance Monte Carlo transport code, which currently does not offer any sensitivity capabilities with MG particle transport simulations. Initial implementation of the new MG TSUNAMI-3D sequence and its preliminary results with a selected critical benchmark experiment in the Verified, Archived Library of Inputs and Data (VALID) are presented in this study.
KENO↗
Sensitivity Calculations for Systems with Polyethylene Reflector Materials Using CLUTCH [Slides]
CLUTCH is a CE TSUNAMI method that uses a single forward calculation to determine sensitivities. An F*(r) function provides the importance of each voxel in a mesh over all regions where fission can occur. A sufficient number of fissions must be simulated in each voxel in which fission is possible to generate an accurate estimate of the importance of a fission in that voxel.
73 NUCLEAR PHYSICS AND RADIATION PHYSICS↗
Adapting CLUTCH Methodology to Multigroup TSUNAMI-3D for Eigenvalue Sensitivity Calculations [Slides]
Abstract not provided.
97 MATHEMATICS AND COMPUTING↗
Method for use of hydraulically or electrically controlled solenoids under failed on conditions
A method to operate a clutch device in an electro-mechanical transmission mechanically-operatively coupled to an internal combustion engine and at least one electric machine includes, in response to a failure condition detected within a flow control device configured to facilitate flow of hydraulic fluid for operating the clutch device, selectively preventing the flow of hydraulic fluid from entering the flow control device and feeding the clutch device. Synchronization of the clutch device is initiated when the clutch device is intended for activation, and only if the clutch device is synchronized, the flow of hydraulic fluid is selectively permitted to enter the flow control device to activate the clutch device.
42 ENGINEERING↗
Multiple Paternity Benefits Female Marbled Salamanders by Increasing Survival of Progeny to Metamorphosis
Abstract Multiple paternity occurs in most species and animal groups that have been studied. Because mating involves fitness costs to individual females, theory predicts that polyandrous females gain greater fitness benefits than costs, allowing the behavior to be maintained. Genetic, rather than material, benefits often occur in species where males provide females with little more than sperm and seminal fluid. We compared fitness correlates of single‐ and double‐sire clutches from female marbled salamanders ( Ambystoma opacum ) at the egg, hatchling, and metamorph stages of offspring development. Because clutches were collected from experimental breeding groups, strict paternity exclusion of offspring using microsatellite data allowed us to categorize each clutch as having either one or two fathers. Early offspring viability and size of hatchlings were not different between single‐ and multiple‐paternity clutches. Larvae from the two clutch types were allowed to develop together in field enclosures until metamorphosis. Although there was no difference in size at metamorphosis, survival to metamorphosis was significantly higher in multiple‐paternity clutches (44% vs. 40%) suggesting a benefit for females. The results were consistent with genetic benefits, although maternal effects could not be ruled out. The data did not support predictions of the genetic bet‐hedging and good sperm hypotheses for genetic benefits of polyandry.
Croshaw, Dean A.↗
Environmental Impacts of Wind Power Development on the Population Biology of Greater Prairie-Chickens
1. We investigated the impacts of wind power development on the demography, movements, and population genetics of Greater Prairie-Chickens (Tympanuchus cupido) at three sites in northcentral and eastern Kansas for a 7-year period. Only 1 of 3 sites was developed for wind power, the 201MW Meridan Way Wind Power Facility at the Smoky Hills site in northcentral Kansas. Our project report is based on population data for prairie chickens collected during a 2-year preconstruction period (2007-2008), a 3-year postconstruction period (2009-2011) and one final year of lek surveys (2012). Where relevant, we present preconstruction data from our field studies at reference sites in the northern Flint Hills (2007-2009) and southern Flint Hills (2006-2008). 2. We addressed seven potential impacts of wind power development on prairie chickens: lek attendance, mating behavior, use of breeding habitat, fecundity rates, natal dispersal, survival rates, and population numbers. Our analyses of pre- and postconstruction impacts are based on an analysis of covariance design where we modeled population performance as a function of treatment period, distance to eventual or actual site of the nearest wind turbine, and the interaction of these factors. Our demographic and movement data from the 6-year study period at the Smoky Hills site included 23 lek sites, 251 radio-marked females monitored for 287 bird-years, and 264 nesting attempts. Our genetic data were based on genotypes of 1,760 females, males and chicks that were screened with a set of 27 microsatellite markers that were optimized in the lab. 3. In our analyses of lek attendance, the annual probability of lek persistence during the preconstruction period was ~0.9. During the postconstruction period, distance to nearest turbine did not have a significant effect on the probability of lek persistence. However, the probability of lek persistence increased from 0.69 at 0 m to 0.89 at 30 km from turbines, and most abandoned lek sites were located <5 km from turbines. Probability of lek persistence was significantly related to habitat and number of males. Leks had a higher probability of persistence in grasslands than agricultural fields, and increased from ~0.2 for leks of 5 males, to >0.9 for leks of 10 or more males. Large leks in grasslands should be a higher priority for conservation. Overall, wind power development had a weak effect on the annual probability of lek persistence. 3. We used molecular methods to investigate the mating behavior of prairie chickens. The prevailing view for lek-mating grouse is that females mate once to fertilize the clutch and that conspecific nest parasitism is rare. We found evidence that females mate multiple times to fertilize the clutch (8-18% of broods, 4-38% of chicks) and will parasitize nests of other females during egg-laying (~17% of nests). Variable rates of parentage were highest in the fragmented landscapes at the Smoky Hills field site, and were lower at the Flint Hills field site. Comparisons of the pre- and postconstruction periods showed that wind energy development did not affect the mating behaviors of prairie chickens. 4. We examined use of breeding habitats by radio-marked females and conducted separate analyses for nest site selection, and movements of females not attending nests or broods. The landscape was a mix of native prairie and agricultural habitats, and nest site selection was not random because females preferred to nest in grasslands. Nests tended to be closer to turbines during the postconstruction period and there was no evidence of behavioral avoidance of turbines by females during nest site selection. Movements of females not attending nests or broods showed that females crossed the site of the wind power development at higher rates during the preconstruction period (20%) than the postconstruction period (11%), and that movements away from turbines were more frequent during the postconstruction period. Thus, wind power development appears to affect movements in breeding habitats but not nest site selection of female prairie chickens during the breeding season. 5. We tested the effects of wind power development on five components of female fecundity: timing of clutch initiation, clutch size of first nests and renests, nest survival, and hatchability of eggs. Average date of clutch initiation was 26 April, clutch size was 12.7 and 10.6 eggs for first nests and renests, probability of nest survival was low at 0.18, but egg hatchability was high at 0.79. Wind power development had no impact on reproductive effort or nesting success, and all five components of fecundity were not related to treatment period or distance to turbine. Nest survival was the main factor limiting reproductive output of female prairie chickens and most losses were due to predation. Daily nest survival was strongly related to vegetative cover at the nest. Changes to rangeland management practices that would double nesting cover from 2.5 to 5 dm would triple the probability of nest survival from 0.17 to 0.52. Grass and forb cover had weak positive effects on daily nest survival whereas shrub cover, proximity to woodlands, and recent rainfall had negative effects. Reproductive performance of prairie chickens is low in managed rangelands in northcentral Kansas and efforts to improve range conditions and reduce predator activity would aid recovery of prairie chicken populations. 6. We used molecular methods to investigate patterns of natal dispersal in prairie chickens. High rates of nest failure limited the number of young that we could sample. Direct detections of natal dispersal were limited because survival of newly hatched chicks to become adults were low and because we were unable to detect dispersal distances outside of our study area. Direct observations of natal movements were limited and were inadequate to make conclusions about the potential impacts of wind energy development on natal dispersal. Spatial correlograms of genetic distance among males at leks were a more sensitive measure of population structure, and indicated a weak effect of wind energy development on the spatial genetic structure of prairie chickens. 7. We tested the effects of wind power development on female survival with time-to-event models, and on residual body mass of males with analyses of covariance. Distance to turbine and the interaction of distance and treatment period had no effect on female survival. Contrary to predictions of negative impacts of wind power development, the probability of female was lowest during the preconstruction period (0.274) and increased significantly during the postconstruction period (0.543). Inspection of hazard functions indicated that the difference in annual survival could be attributed to a higher risk of mortality during the lekking season in the preconstruction period. We suggest that wind power development may have improved ecological conditions for prairie chickens by disrupting the foraging behavior of diurnal raptors that kill prairie chickens at lek sites. In support of this idea, raptor kills tended to be farther from turbines during the postconstruction period whereas mammalian kills were closer. Analyses of the major causes of mortality did not support our hypothesis because the odds of raptor predation were greater after development was completed. Most mortality losses of radio-marked females were due to predation, and losses to collision mortality or harvest were rare events. Low rates of natural mortality during fall and winter imply that harvest is likely to be additive mortality in prairie chickens. Wind energy development reduced the residual body mass of male Greater Prairie-Chickens at lek sites near turbines. Low values of residual body mass could have a negative impact on individual survival or fecundity rates, or may be related to predation risk and flight performance of males displaying at open lek sites. 8. We tested for impacts of wind power development on population numbers of prairie chickens with monitoring of male numbers at leks, and with genetic measures of population structure. Lek counts indicated that wind power development did not affect the population size of prairie chickens. Peak counts of males at leks were recorded the first year after construction was completed and the highest rates of population change were observed during the interval when the wind power facility was constructed. Population numbers of prairie chickens near and distant from turbines appeared to covary in parallel, probably because bird numbers were linked by dispersal movements. Estimates of population viability based on genetic diversity, effective population size and rates of population exchange did not show annual changes and were unaffected by wind development during our study. Estimates of relatedness among males at the same and different leks suggested that wind power development has either reduced dispersal rates or changed settlement patterns, leading to higher rates of relatedness among males displaying at the same lek site. 9. Greater Prairie-Chickens were not strongly affected by wind power development in Kansas. Negative impacts of wind power development included a trend for reductions in lek persistence near turbines, behavioral avoidance of turbines by females during their breeding season movements, and changes in the genetic structure of males at leks that were consistent with reduced dispersal or recruitment rates. We found no impacts of wind power development on nest site selection, female reproductive effort or nesting success, or population numbers. Positive impacts of wind power development included an increase in female survival rates. We hypothesized that the unexpected increase in female survival was related to changes in trophic interactions and disruption of the foraging behavior of raptors that kill prairie chickens at lek site. 10. Research funding for this project included a grant from the 20% Wind by 2030 Program of the Department of Energy (this final report), and grants from the Kansas Department of Wildlife, Parks and Tourism, the National Fish and Wildlife Federation, and initial funding from the National Wind Coordinating Collaborative. Research products from data collected during the preconstruction period have included five peer-reviewed research articles and two PhD dissertations at Kansas State University. Additional manuscripts are in review for possible publication in 2013-2014.
17 WIND ENERGY↗
Control system for a hybrid powertrain system
A vehicle includes a powertrain with an engine, first and second torque machines, and a hybrid transmission. A method for operating the vehicle includes operating the engine in an unfueled state, releasing an off-going clutch which when engaged effects operation of the hybrid transmission in a first continuously variable mode, and applying a friction braking torque to a wheel of the vehicle to compensate for an increase in an output torque of the hybrid transmission resulting from releasing the off-going clutch. Subsequent to releasing the off-going clutch, an oncoming clutch which when engaged effects operation of the hybrid transmission in a second continuously variable mode is synchronized. Subsequent to synchronization of the oncoming clutch, the oncoming clutch is engaged.
33 ADVANCED PROPULSION SYSTEMS↗
Deterministic-Monte Carlo Hybrid Methods for Eigenvalue Sensitivity Coefficient Calculations
The TSUNAMI suite within the SCALE code package includes several methods for generating sensitivity data, including multigroup (MG) and continuous-energy (CE) capabilities. For generating sensitivities with CE data, three methods are available in SCALE 6.3.0: (1) the iterated fission probability (IFP) method with the KENO Monte Carlo transport solver, (2) IFP with the Shift Monte Carlo transport solver, and (3) the Contributon-Linked eigenvalue sensitivity/Uncertainty estimation via Tracklength importance Characterization (CLUTCH) with the KENO Monte Carlo transport solver. Currently, it is difficult to generate accurate sensitivities with large reflectors when using the CLUTCH method, specifically with fissionable and hydrogenous materials. To address this issue, the work presented herein examines a methodology to calculate the adjoint flux externally with the 3D deterministic SN transport code DENOVO in SCALE; the result is then read directly into the CLUTCH-TSUNAMI sequence. This hybridization method replaces the Monte Carlo F*(r) calculation in CLUTCH while still utilizing the forward calculation. The critical benchmark HEU-MET-FAST-028-001 is used to generate sensitivities based on the inability of CLUTCH to generate accurate sensitivities. Results from the hybrid method appear to generate sensitivity values that are in excellent agreement with direct perturbations. Although further testing is needed, the method provides promising results for the development and utility of a hybrid method for use in TSUNAMI.
73 NUCLEAR PHYSICS AND RADIATION PHYSICS↗
Analysis of SCALE Criticality and Sensitivity Calculations for Reflected HEU Cylinders
The SCALE code package offers multiple nuclear data libraries and sensitivity and uncertainty (S/U) methods supporting and derived from Monte Carlo (MC) transport. The CSAS and TSUNAMI-3D sequences use KENO MC, utilizing either continuous-energy (CE) cross sections or multigroup (MG) cross section libraries. TSUNAMI-3D has two CE calculational methods: the iterated fission probability (IFP) method, and the Contribution-Linked eigenvalue sensitivity/Uncertainty estimation via Tracklength importance CHaracterization (CLUTCH) method. Previous work has shown poor agreement between CLUTCH and confirmatory direct perturbation calculations in specific applications (e.g., fissionable and polyethylene reflectors). The HEU-MET-FAST-084 (HMF-084) International Criticality Safety Benchmark Evaluation Project evaluation consists of 27 cylindrical highly enriched uranium metal cores with 14 unique reflector materials of 0.5 and 1 in. thicknesses. Included in this list of reflector materials are natural uranium and polyethylene. This work utilized SCALE 6.2.4 models of the HMF-084 evaluation, with additional non-physical configurations to test both the MG bias and CLUTCH functionality across a variety of reflector material thicknesses. The evaluation’s use of concentric cylinders allowed for examination of several MG self-shielding methods: infinite homogenous, cylindrical, and spherical. The results indicate that the use of polyethylene reflectors with CLUTCH is not fundamentally impossible but sensitive to geometry. The poor performance of CLUTCH with fissionable reflectors was reaffirmed. The 2 in. and greater polyethylene-reflected calculations demonstrate the necessity of using the 302-group library for fast systems. The nickel MG bias was substantial, as discussed in a companion paper, as were cobalt and iron.
73 NUCLEAR PHYSICS AND RADIATION PHYSICS↗